Simulate a structured population with temporal autocorrelation using standard Leslie matrices. Each element in the Leslie matrix has a specified mean, variance, and temporal autocorrelation value. The matrix can have arbitrary dimensions and can have transitions besides linear survival. This model includes environmental stochasticity with colored noise. Density dependence and demographic stochasticity not currently supported.

matrix_model(data, initialPop, timesteps, covMatrix = NULL, colNames = NULL,
  matrixStructure = NULL, repeatElements = NULL,
  survivalOverflow = "scale")

Arguments

data

The input data can be one of two formats: a list of three matrices, or a data frame with three columns.
If it is a list of three matrices, they must be standard Leslie matrices: the first a matrix of mean values for each matrix element, the second a matrix of standard deviations, and the third a matrix of temporal autocorrelations.
If it is a data frame, there must be three columns, one for mean vital rates, one for standard deviations, and one labeled 'autocorrelation.'
If the population has n stages, the first n rows of the data frame must be the matrix elements for the first stage, and the next n*(1-n) rows must be the transition probabilities, each row of the matrix from first to last transposed vertically.
If you want to run a matrix population model without temporal autocorrelation, simply set all autocorrelation values to zero.

initialPop

An initial population vector. The length must be the same as the number of classes in the matrices.

timesteps

The number of timesteps you would like to simulate the population.

covMatrix

Optional: Add a covariance matrix describing within-year covariances between matrix elements. The matrix elements must be in the same order as they are in the data frame format above: a Leslie matrix turned into a vector row-wise. There should be as many columns as matrix elements, excluding repeat elements (see below) or structural zeros.

colNames

Optional: If the mean, sd, and autocorrelation columns of your data frame input are not named 'mean', 'sd', and 'autocorrelation', provide their names here in a character vector, e.g., `c(mean = 'Mean', sd = 'Standard Deviation', autocorrelation = 'phi')`

matrixStructure

Optional: By default, the function assumes that the first row of the matrix gives fecundities while the rest of the matrix gives transition or survival probabilities. However, these assumptions do not apply to many plant matrices. If your matrix has transition probabilities in the first row or fecundities beyond the first row (e.g., clonal reproduction), provide a character matrix here with the same dimensions as your matrix that gives in strings whether each element is 'fecundity' or 'transition'.

repeatElements

Optional: Sometimes not all matrix elements can be measured, and some transitions or fertilities are generalized across classes. If you have any matrix elements that are copies of other matrix elements (e.g., stage 3 is assumed to have the same fertility as stage 4) indicate them here with a matrix of rowwise (not column-wise) indices that show which elements are repeats and which are unique. For example in a 2x2 matrix where both classes are assumed to have the same fertility, input `matrix(c(1, 1, 3, 4), byrow = T, ncol = 2)`. If you indicate repeat elements and you include a covariance matrix, the covariance matrix must only have as many columns as unique matrix elements. Structural zeros should not be included here as repeats, as they are automatically detected in the function.

survivalOverflow

If the survival for a stage is very high or very variable, the function may sometimes generate projection matrices with survival that exceeds 1 for that stage. The function has two methods of dealing with this problem: either discard all projection matrices and generate new ones until the survival falls within acceptable bounds ("redraw") or divide all the non-fertility matrix elements for that stage by the survival such that they add to 1 ("scale"). The default is "scale".

Value

A data frame with n + 2 columns, where n is the number of stages in the matrix. One column indicates the timestep, there is one column with the population size for each stage, and one column for total population size.

Examples

meanMat <- matrix(c(0.55, 0.6, 0.24, 0.4), byrow = TRUE, ncol = 2) sdMat <- matrix(c(0.3, 0.35, 0.05, 0.1), byrow = TRUE, ncol = 2) phiMat <- matrix(c(-0.2, -0.2, 0, 0), byrow = TRUE, ncol = 2) initialPop <- c(100, 100) sim <- matrix_model(list(meanMat, sdMat, phiMat), initialPop, 50) head(sim)
#> # A tibble: 6 x 4 #> timestep total stage1 stage2 #> <int> <dbl> <dbl> <dbl> #> 1 1 200 100 100 #> 2 2 191. 98.8 91.9 #> 3 3 200. 53.9 146. #> 4 4 154. 77.1 77.2 #> 5 5 126. 57.9 68.0 #> 6 6 95.1 48.4 46.7